Run to Seed by Phi

Puzzle explanation

The thematic unclued across entries represented a salmon ladder, a series of steps to help salmon find their way upstream around obstacles. They were clued in pairs from the bottom of the “ladder” upwards, from SALMON to LADDER with five types of salmon in between. The unclued down entry was OUANANICHE, another type of salmon. The title referred to a salmon run, the migration of salmon up a river from the sea, in order to spawn.

Clue explanations

Conventions: * = anagram, < = reversal

No Jumbles Answer Explanation
Double
  SALMON, GRILSE LISSOM L + IS + SOM(e)
ANGLER ANGLE + R
  GRILSE, LIGGER LEGGIER (G + I) in LEGER
GIRLS (S + L + RIG) <
  LIGGER, SAMLET LARGE L + R in AGE
GIMLETS GETS around (I + ML)
  SAMLET, BAGGIT BIGGEST GIG * in BEST
MALTA A after MALT
  BAGGIT, KIPPER GRIPPE GRIPPE(d)
KITBAG K(ill) + IT + GAB <
  KIPPER, LADDER RIPPLED L in RIPPED
DRAKE RAKE after D
Across
1   SOFFIT SO + F + FIT
9   QUALIA QU + (A + A) around LI
10   EMERGE (EG + REME) <
11   LARES L + ARES
13   ONE-STEP (HONEST − H) + EP
14   AYRE AYRSHIRE − SHIR
17   ENSUE ((E + USE) around N) <
18   JENNET JET around (NINE − I)
23   ACHAEA (ACHE + A) around A
25   EMEER E’ER around ME
27   DEEV VE(tt)ED <
29   AVIETTE A + VIGNETTE − (G + N)
31   ORRIS OR + RISK − K
32   ELYSÉE ELY + SEE
33   SIMEON E in SIMON
34   TRIENE TRIE(s) around EN
Down
2   FAR EAST FT around AREAS
3   FLESH SELF < + H
4   TABETIC ABET in TIC(k)
5   LEI LEIR − R
6   CÉZANNE (ENC around NAZE) <
7   ARMY BARMY − B
8   REFECTORIES (C + TORIES) after FREE *
11   LOVE HANDLES (SLAVE HELD ON) *
12   SAFE S(ellers) A(ctor) F(rom) E(ngland)
15   RE-ELECTION REEL + (s)ECTION
16   BEA BEA(m)
19   CANVASS VAS in CANS
20   LIEVEST LIVES around E + T
21   E’ER (b)EER
22   YEW TREE YET around W + REE
24   EXAM E + MAX <
26   FERMI REF < + MI
28   ERYL LYRE <
30   SEN 2 meanings
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