## Singles Only by Twin

### Puzzle explanation

The cells containing two letters were at all the crossings of rows and columns 1, 5, 9 and 13 (as shown in the initial grid fill here), and spelt “clear and shade these columns and rows”. In normal order of grid entries, the extra letters in wordplay spelt out “letter’s first position in its clue’s letters”, detailing how to replace the remaining letters with numbers (for all of the checked letters, the position was the same in both the across and the down clue), and “erase ten plus”, so the numbers greater than 9 were also cleared. Letters that weren’t convertible to numbers because they didn’t occur in their clues (shown in blue here) spelt out “complete sudoku”; erasing them left a sudoku puzzle to be solved.

### Clue explanations

Conventions: * = anagram, < = reversal

Answer Explanation [extra letter]
ACUSHLA (BUSH − B + LA) after AC[T]
AD HOC (D + HO[T]) in AC
ALIDADE (I’D [R]EAD) * after (A + L)
AMAIN MA < + [P]AIN
AMIS A + MIS[E]
ANTHER [T]AN + THE + R
ARABIN [O]AR + A + BIN
ASPIRE A[S] around SPIRE
BLUER (c)LUE(s) in (m)[E](m)B(e)R
BRANE B[O]RANE
BRETASCHE (HE ACTS) * in BR[I]E
BUS BUS[S]
CLOTH-EARS CLOT + H + [L]EARS
COLIN COL + [R]IN
CO-OPTS OP[E](n) in COST *
COTES COT[T]AGES − AG
DEO [L]EO around D
DRONGO DIR[T] − I + GOON *
ESTRAY [R]EST DAY with R for (an)D
GIUST GUST around I[I]
GOVERN NOG < around VE[E]R
GRADUAND ARGU[E]D + AND
HERONSEWS HERO[I]N + SEWS
HOP IT [C]HIT around OP
LISLE L + S in [R]ILE
MAN-EATER ANE in MA[S]TER
NEIGH NEI[N] + G + H
ODONTIC (TIN + C(an)) * after [T]O-DO
OEDEMATA (MADE TO EA[T]) *
OPAL O + AL[L] after P(ickling)
PERSUADER (RED + AU + (vest)S (wa)R (pain)[T] (whit)E (ca)P) <
RATCHETED TAR < + CHE[S]TED
RECOMMEND RE + COM[E] + MEND
REMNANT (TRAN[S] MEN) *
RESTEMS (M + [L]ESTER + S) *
SALTANDO SALO[N] around (s)TAND
SCRUB S + U in C[U]RB
SERRICORN (SINC(e) + ERROR[S]) *
SKA SK[U]A
SKEGG SA[N]K − A + EGG
SPASM M[I]SHAPS < − H(ousehold)
TEST TE[F] + ST
THO’ HO[S]T *
TOED (D[I]E + OT) <
TRADE (bar)T[E]R A DE(al)
TRIPEDAL RIP[P]ED in LAT <
TSETSE TE[E] around TESS *
UNITISE (SIT IN) < in U[A]E
UNVIRTUE (E + VIRU[S] NUT) *
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